1 1 2 Receptor usage and cell entry of porcine epidemic diarrhea coronavirus 3 4 Chang Liu 1, #, Jian Tang 2, #, Yuanmei Ma 3, #, Xueya Liang 3, Yang Yang 1, 5 Guiqing Peng 4, Qianqian Qi 2, 5, Shibo Jiang 2, 5, Jianrong Li 3, Lanying Du 2, *, 6 Fang Li 1, * 7 1 Department of Pharmacology, University of Minnesota Medical School, Minneapolis, 8 MN 55455, USA 9 2 Lindsley F. Kimball Research Institute, New York Blood Center, New York, NY 10065, 10 USA 11 3 Department of Veterinary Biosciences, College of Veterinary Medicine, The Ohio State 12 University, Columbus, OH 43210, USA 13 4 State Key Laboratory of Agricultural Microbiology, College of Veterinary Medicine, 14 Huazhong Agricultural University, Wuhan, Hubei, China 15 5 Key Laboratory of Medical Molecular Virology of Ministries of Education and Health, 16 Shanghai Medical College and Institute of Medical Microbiology, Fudan University, 17 Shanghai, China 18 19 20 #These authors contributed equally to this work. 21 22 * Correspondence: 23 Fang Li (