ORIGINAL ARTICLE Identification of a Lineage D Betacoronavirus in Cave Nectar Bats (Eonycteris spelaea) in Singapore and an Overview of Lineage D Reservoir Ecology in SE Asian Bats I. H. Mendenhall1, S. Borthwick1, E. S. Neves1, D. Low1, M. Linster1, B. Liang1, M. Skiles2, J. Jayakumar1, H. Han3, V. Gunalan3, B. P. Y.-H. Lee4,5, K. Okahara1, L.-F. Wang1, S. Maurer-Stroh3,6, Y. C. F. Su1 and G. J. D. Smith1,7 1 Programme in Emerging Infectious Diseases, Duke-NUS Medical School, Singapore 2 College of Veterinary Medicine, North Carolina State University, Raleigh, NC, USA 3 Bioinformatics Institute, Agency for Science, Technology and Research, Singapore 4 Durrell Institute of Conservation and Ecology, School of Anthropology and Conservation, University of Kent, Canterbury, UK 5 National Parks Board, Singapore 6 School of Biological Sciences, Nanyang Technological University, Singapore 7 Duke Global Health Institute, Duke University, Durham, NC, USA Keywords: disease ecology; virus diversity; evolution; distribution Correspondence I. H. Mendenhall and G. J. D. Smith. Programme in Emerging Infectious Diseases, Duke-NUS Medical School, Singapore. Tel.: +65 6601 1109; Fax: +65 6221 2529; E-mails: