Neurovirulent murine coronavirus JHM.SD uses cellular zinc metalloproteases for 1 virus entry and cell-cell fusion 2 3 Running title: Metalloproteases in murine coronavirus fusion 4 5 Judith M. Phillips1#, Tom Gallagher2, and Susan R. Weiss1 6 7 1) Department of Microbiology, Perelman School of Medicine, University of 8 Pennsylvania, Philadelphia, PA, USA 9 2) Department of Microbiology and Immunology, Loyola University Medical Center, 10 Maywood, Illinois, USA 11 12 Address correspondence to: Judith M. Phillips,